This paper reviews research into the hydrological impacts of UK upland forestry and updates the water balance of the Plynlimon research catchments for the period 19722004. Comparison of this network of densely instrumented coniferous forest and grassland catchments builds upon previously reported differences in annual evaporation of the two land uses and, most crucially, provides evidence of systematic, agerelated, variations in forest evaporation losses over a managed plantation forest cycle. In comparison with the grassland catchment, the additional water use of the 70% forested catchment fell from 250 to 150 mm yr 1 because of increasing forest age; this is equivalent to a decline from 370 mm to 210 mm extra evaporation from a complete forest cover. At present, with up to half of the forest area felled or only recently replanted, the difference in evaporation between the forest and grass catchments is negligible. Knowledge of the period of maximum tree water use may be critically important for the future management of multi-use forests. This is also being investigated by micro-meteorological measurements at the scale of the forest stand using eddy covariance, in conjunction with the long-term catchment monitoring.