As an extension of M. Hayes' argument (Hayes, 1977), a general relationship is derived between group velocity and fluxes quadratic in wave amplitude for single, weakly-modulated small amplitude plane waves propagating in an inhomogeneous, non-conservative, dispersive system. If a quadratic (4-) flux constructed from wave quantities with a common single harmonic phase is inserted into its linear governing equation, the resultant equation consists of a wave part with a hi-harmonic phase and a mean part, but both parts must vanish separately to satisfy the equation. The amplitude of the former gives the dispersion relation, H(k*)=0; this in turn gives the complex (4-)group velocity vector *g=*H/ * by varying the complex (4-)wavenumber vector k*(*=0, 1, 2 and 3). On the other hand, the latter gives a governing equation for the mean of the quadratic flux, and also gives, by taking the same variation, another mean quadratic flux. The latter flux is proportional to the group velocity *g. The general relationship is illustrated by application to Rossby wave motions. The group velocity and energy flux relation obtained by Longuet-Higgins (1964) is rederived as a special case.