“…Capture-recapture techniques for estimating various population parameters include physical tagging (Baker, 2004), genetic tagging (Palsbøll et al, 1997) or photographic identification (photo-ID) of individuals through their natural markings (Würsig and Jefferson, 1990;Hammond, 2009), including pigmentation patterns (Arzoumanian et al, 2005), pelage marks (Hastings et al, 2008), scars (Gilkinson et al, 2007), and whisker patterns (Anderson et al, 2010). Photo-ID has been widely used across taxonomic groups, such as terrestrial and marine mammals (Kelly, 2001;Langtimm et al, 2004), reptiles (Schofield et al, 2008;Gardiner et al, 2014) and fish (Couturier et al, 2014;Kanive et al, 2015).…”