“…Model simulations indicate an upward shift of the lower limit of their range in the Rif Mountains by about 200 m (Cheddadi et al, 2017). Recent decline in these mountains, however, is considered to have been induced by human activities, mainly cropping, cutting and overgrazing (Ajbilou et al, 2003; Benabid, 1985; Coudel et al, 2016; Grovel, 1996; Hamelin and Nwankwo, 2013; Moore et al, 1998; Rejdali, 2004). Disentangling the impact of each factor on landscape is not an easy task, although essential to take the appropriate managing measures to ensure the persistence of the modern populations (Allen et al, 2010; El-Baha et al, 2010; IRES, 2010; Vegas-Vilarrúbia et al, 2011).…”