The Himalayan foreland basin and other parts of the world had witnessed a dramatic change in the ecological structure during the Late Miocene, as indicated by the increase in abundance of C 4 plants in a C 3 dominated ecosystem. However, the asynchronous expansion of C 4 plants across the latitudes, as well as within the different sub-basins of the Himalayan foreland basin, tentatively suggests that regional climatic factors and sedimentary architecture vis-à-vis geomorphological settings might have an important influence in controlling the abundance of C 4 plants. This study has carried out sedimentological and palaeohydrological analyses of the Siwalik channels at Naladkhad and Ranital regions of the Kangra sub-basin, north-west Himalaya, and estimated slope and sinuosity of the modern Himalayan fan rivers (n > 100), which provides a hydrological analogue for the Siwalik channels. The results suggest that the Siwalik Group in the Kangra sub-basin was deposited by braided rivers, situated at the proximal part of palaeo-alluvial fan. In addition to climate-driven changes, the comparison between sedimentary architecture and published carbon isotope data from the Kangra sub-basin suggests that channel-fill dominated fan-proximal Naladkhad and Ranital regions favoured the growth of C 3 plants until ca 6 Ma, a time when Pakistan Siwaliks were dominated by C 4 plants. In comparison to the Kangra sub-basin, Pakistan Siwaliks are characterized by a higher abundance of floodplain sediments and possibly represent a distal fan deposit. Data from other parts of the world similarly suggest that, in addition to climate forcing, the dominance of overbank fines favoured the growth of C 4 plants. Therefore, detailed knowledge of the depositional environment and palaeo-geomorphic setting of the sedimentary archives is essential to understand the influence of sedimentary architecture on the spatio-temporal variation in the abundance of C 4 vegetation, especially for foreland basin settings, where lateral facies transition rate is high.