“…However, studying elasmobranch dentitions is not an easy task because of various intraspecific variation parameters one must take into consideration. Intraspecific tooth variations in elasmobranchs can occur in number of tooth rows, in number of accessory cusps (e.g., Gudger, 1937;Sadowsky, 1970;Taniuchi, 1970), at different ontogenetic stages (e.g., Reif, 1976;Shimada, 2002b); between sexes that may also differ between different mating seasons (e.g., McCourt and Kerstitch, 1980;Gruber and Compagno, 1981;Kajiura and Tricas, 1996), among different geographic regions (Lucifora et al, 2003), or in abnormal forms (e.g., Gudger, 1937;Raschi et al, 1982;Kaneko and Goto, 2001). The lack of any consideration of such variations would lead to the 'taxonomic instability' of toothbased species common to fossil forms (Shimada and Cicimurri, 2006).…”