“…When investigating the biofilm thickness in a trickling filter, Persson et al (2002) effectively found that it decreases with depth. Moreover, potential nitrifying and denitrifying activities measured ex-situ were much greater than activities observed inside the trickling biofilters, suggesting the possibility to enhance nitrogen removal (Gilbert et al, 2007(Gilbert et al, , 2008. Moreover, potential nitrifying and denitrifying activities measured ex-situ were much greater than activities observed inside the trickling biofilters, suggesting the possibility to enhance nitrogen removal (Gilbert et al, 2007(Gilbert et al, , 2008.…”