“…Red tides -discolorations of the sea surface due to dense plankton blooms -occur along much of the world's coastline and in offshore waters (Holmes et al, 1967;Eppley and Harrison, 1975;ECOHAB, 1995;Jeong, 1995;Horner et al, 1997;Imai et al, 2001;Sordo et al, 2001;Anderson et al, 2002;Jeong et al, 2003Jeong et al, , 2008Jeong et al, , 2013bAlonso-Rodriguez and Ochoa, 2004;Seong et al, 2006;Johnson et al, 2013;Kang et al, 2013;Lee et al, 2013c;Park et al, 2013a). Red tides frequently cause large-scale mortalities of fish and shellfish and significant losses to the aquaculture and tourist industries of many countries (Shumway, 1990;Smayda, 1990;Shumway and Cembella, 1993;Glibert et al, 2005;Anderson et al, 2012;Fu et al, 2012;Park et al, 2013d).…”