“…Human impacts (e.g., persecution, disturbance) result in different behavioural responses of large carnivores, such as wolves, to minimise their interactions with humans and their effects (e.g., Whittington et al, 2005;Llaneza et al, 2012;Lesmerises et al, 2013;Ahmadi et al, 2014). Factors influencing exposure risk, such as the availability of refuge habitat, are also modulated by human activity and impact at the landscape level (Thurber et al, 1994;Llaneza et al, 2012).…”