This research was conducted under the auspices of the graduate school for Production Ecology and Resource Conservation (PE&RC). Chapter 2 Effects of large herbivores on wood pasture dynamics in a European wetland system Cornelissen, P., Bokdam, J., Sýkora, K., Berendse, F. (Basic and Applied Ecology 2014, 15, 396-406.) Chapter 3 Transition of a Sambucus nigra L. dominated woody vegetation into grassland by a multi-species herbivore assemblage Cornelissen, P., Gresnigt, M., Vermeulen, R., Bokdam, J., Smit, R. (Journal for Nature Conservation 2014, 22, 84-92.) Chapter 4 Effects of floodplain restoration and grazing on wood encroachment along a lowland river in NW-Europe Cornelissen, P., Decuyper, M., Sýkora, K., Bokdam, J., Berendse, F. (Submitted) Chapter 5 Density dependent diet selection and body condition of cattle and horses in heterogeneous landscapes Cornelissen, P., Vulink, J.Th. (Applied Animal Behaviour Science 2015, 163, 28-38.) Chapter 6 Rewilding Europe: Early dynamics of a multispecies grazing ecosystem Cornelissen, P., Vera, F.W.M., Berendse, F., Sýkora, K., Bokdam, J., Ritchie, M.E., Olff, H. Chapter 7 Effects of weather variability and geese on population dynamics of large herbivores creating opportunities for wood-pasture cycles. A modelling approach Kramer, K., Cornelissen, P., Groot Bruinderink, G.W.T.A., Kuiters, L., Lammertsma, D., Vulink, J.Th., Van Wieren, S.E., Prins, H.H.T. Chapter 8
GENERAL INTRODUCTIONScope of this study Conservation and restoration of biodiversity are major objectives for managers of nature reserves and in many cases large herbivores are considered to be 'keystone' species to achieve these goals (e.g. Wallis De Vries et al. 1998;Zabel and Anthony 2003;Danell et al. 2006;. Large herbivores are major drivers of changes in the structure and functioning of terrestrial ecosystems, because they modify nutrient cycles, soil properties, net primary production, patterns in vegetation structure and composition, and fire regimes (Gordon 2006). Through their impact on plant community structure (openness and height) (e.g. Thompson Hobbs 2006;Smit and Putman 2011), large herbivores affect most other plant and animal species in these communities (e.g. Root 1973;Cody 1975;Suominen and Danell 2006). To restore and maintain species diversity, controlled grazing with large wild and domestic herbivores at low stocking rates has developed towards a major strategy for conservation management in Europe (e.g. Wells 1965;Gordon et al. 1990;. In this context, traditional livestock farming landscapes (e.g. man-made wood pastures, heathland, chalk grasslands) serve as a reference (Pott and Hüppe 1991;Bignal et al. 1994;Piek 1998). The strategy of controlled grazing has been challenged by a more 'natural' grazing strategy, which is inspired by present natural or near-natural grazing systems in Africa and North America, but also by grazed ecosystems in the remote past, which were present during different Pleistocene and Holocene periods (e.g. Van de Veen 1975; Soulé and Noss ...