“…Anthocyanins are synthesized in cytosol, and stored in the vacuole. Studies on several plant species, including Arabidopsis ( Baudry et al, 2006 ; Gonzalez et al, 2008 ; Qi et al, 2011 ; Xie et al, 2016 ), agricultural crops ( Yang et al, 2019 ; Dong et al, 2020 ), fruits ( Rahim et al, 2014 ; Zhou et al, 2015 ; Jiang et al, 2019 ; Li C. et al, 2020 ), vegetable and ornamental plants ( Suzuki et al, 2016 ; Xu et al, 2017 ; Jin et al, 2018 , 2019 ; Zhu et al, 2019 ) have revealed that biosynthesis of anthocyanins are controlled by structural and regulatory genes that take part in formation as well as regulation of specific enzymes. The key enzymes including phenylalanine ammonia lyase (PAL), cinnamic acid 4-hydroxylase (C4H), 4-coumarate-CoA ligase (4CL), chalcone synthase (CHS), chalcone isomerase (CHI), flavonone 3-hydroxylase (F3H), dihydroflavonol 4-reductase (DFR), flavonoid 3′-monooxygenase (F3′H), anthocyanin synthase (ANS), as well as UDP-glucose-flavonoid 3- O -glucosyltrasnferase (UFGT) are important in anthocyanin biosynthesis ( Koes et al, 2005 ; Li et al, 2018 ).…”