“…5 , Box 1 ). Additional Hi-C data from a plethora of filamentous fungi show that the centromeres contact independent of—and distinct from—telomere clustering and thus confirmed the existence of Rabl chromosomes, including in ascomycetes [ Penicillium oxalicum (Li et al 2022 ), E. festucae (Winter et al 2018 ), V. dahliae (Seidl et al 2020 , Torres et al 2023 ), Candida albicans (Burrack et al 2016 ), Fusarium verticillioides (Yao et al 2023 ), and Cladosporium fulvum (Zaccaron et al 2022 )] and basidiomycetes [ Puccinia polysora (Liang et al 2022 ), Puccinia graminis (Sperschneider et al 2021 , Henningsen et al 2022 ), Austropuccinia psidii (Edwards et al 2022 ), and Agaricus bisporus (Hoencamp et al 2021 )] ( Table S1 , Supporting Information ). Interestingly, R. irregularis does not show clear centromere bundling for organizing its chromosomes, but it does seem to exhibit telomere bundling (Yildirir et al 2022 ).…”