“…Moreover, they have been widely developed and used in the study of population genetics of many species (Guo et al, 2016;Mohanty et al, 2016;Tan et al, 2016). Furthermore, some of these microsatellite markers developed for Scomber species have been successfully used for amplification in other related species (Tang et al, 2009;Divya et al, 2016). However, it is not very sufficient, and some locus-by-population tests exhibited significant departure from Hardy-Weinberg equilibrium (HWE) (Cheng et al, 2015).…”