“…Physicochemical properties of the recipient environment, such as surface area, depth, substrate, Ca concentration, pH, temperature, and trophic status, have been suggested as main factors constraining density, growth, and invasion success of zebra mussels at regional (among-lake) scales (Ramcharan et al 1992b, Mellina and Rasmussen 1994, Sprung 1995, Naddafi 2007, Whittier et al 2008, whereas biotic factors, such as fish predation, have been suggested as secondary factors (Thorp et al 1998). McMahon (1991) proposed fishes as the most active predators of settled zebra mussels, but only a handful of studies have indicated the potential of fish predation to control invasion success and density of exotic mussels (Eggleton et al 2004, Bartsch et al 2005, Watzin et al 2008.…”