“…A key limitation of these models, however, is that they result in uniformly distributed patterns of space use, meaning encounters between individuals also occur uniformly in space. In stark contrast, most real animals exhibit non-uniform space use within spatially restricted home ranges (Burt, 1943;Bowen, 1982;Powell, 2000;Moorcroft et al, 2006b;Kie et al, 2010;Fleming et al, 2014a;Noonan et al, 2019b;Martinez-Garcia et al, 2020), and encounters between individuals do not occur uniformly in space, but are instead concentrated at territorial boundaries (Nievergelt et al, 1998;Bermejo, 2004;Wilson et al, 2012;Ellwood et al, 2017), in/around heavily used habitats and/or habitat-features (Weckel et al, 2006;Whittington et al, 2011), or at key resources (De Boer et al, 2010;Price-Rees et al, 2013). We therefore base our estimation framework on recent analytical work by Martinez-Garcia et al (2020) incorporating non-uniform movement within home ranges into encounter theory.…”