“…We used a simple massbalance model to estimate the total ecosystem N storage and speculated that soil, tissue, and litter pools would begin to exhibit significant N enrichment as soon as cumulative N addition exceeded storage capacity. We used well-accepted indicators of N enrichment, including soil pH and the tissue and litter N concentration and the C:N ratio, which have been used to estimate the critical load for N in a variety of ecosystems (Cresser, 2000;Fenn et al, 2008Fenn et al, , 2010Gundersen et al, 1998;Horswill et al, 2008;Skeffington et al, 2007;Wilson et al, 2009). Here we report the potential N uptake capacity for these semi-arid shrublands and the cumulative N input that results in significant N enrichment of soil, tissue, and litter pools.…”