“…Only 2 species are shared among all 4 sites: Centrocercus urophasianus and Bubo virginianus. Sage-grouse occur only where extensive stands of sagebrush are present in valley bottoms (Johnsgard 1973) and are thus a good indicator species for that habitat, past and present. Great Horned Owls occupy a diversity of habitats, from tundra to coniferous forests, riparian habitats, and valley bottoms (de la Torre 1990), so the species' presence in all of the cave sites is not surprising, especially given its predilection for caves and cliffs for nesting and roosting.…”