“…Such multistep oxidations are common for oxygenases (Bühler and Schmid, 2004; Guengerich et al, 2011) and examples include the oxidation of amorpha‐4,11‐diene to artemisinic acid by CYP71AV1 (Ro et al, 2006) and of ent ‐kaurene to ent ‐kaurenoic acid by CYP701A3 (Helliwell et al, 1999). Perillyl alcohol oxidation may also be catalyzed by strain‐intrinsic enzymes, as it was reported, for example, for the geraniol dehydrogenase of Castellaniella defragrans (Lüddeke et al, 2012). Several P. putida strains have been reported to convert limonene to perillic acid, for example, P. putida PL (Dhavalikar and Bhattacharyya, 1966), P. putida GS1 (Speelmans et al, 1998), and P. putida DSM12264 (Mirata et al, 2009).…”