“…This same approach was used in some of the work that showed that chimpanzees (and bonobos) are our closest living relatives. 24,25 P EA , P EM , and P AM are similar to previously defined admixturequantifying statistics, such as D-statistics, 23 E-statistics, 12 enhanced D-statistics, 26 and the D 4P statistic. 11 All of these are counting statistics that condition on the presence or absence of derived alleles across individuals.…”