“…A number of different updating mechanisms can be used to determine the evolving state of the graph, specifying how the composition of the population changes under natural selection. A considerable body of research has been devoted to this subject (Abramson and Kuperman, 2001;Santos and Pacheco, 2005;Antal et al, 2006;Ohtsuki et al, 2006;Ohtsuki and Nowak, 2006a,b;Pacheco et al, 2006;Santos et al, 2006a,b;Szabó and Fáth, 2007;Taylor et al, 2007;Fu and Wang, 2008;Roca et al, 2009b;Tarnita et al, 2009a,b;Perc and Szolnoki, 2010;Fehl et al, 2011;Allen et al, 2012;Cavaliere et al, 2012). Remarkably, Ohtsuki et al (2006) derived a simple rule as a good approximation for general graphs according to which natural selection favors cooperation if the benefit of the altruistic act, b, divided by its cost, c, exceeds the average number of neighbors, k (i.e., b=c 4 k implies cooperation).…”