“…Furthermore, he supported the view that Tungusic vowel harmony is retracted tongue root (RTR) based, as it is in Manchu and Evenki, and that the reduced vowel harmony in Contemporary Korean derives from a tongue root based system in Middle Korean. As far as the harmony-like opposition between o 2 and u, o 1 or a in Old Japanese is concerned, the recent reconstruction of a seven-vowel system in Proto-Japonic by Frellesvig and Whitman (2008) implies an underlying opposition between pJ *ɨ, *ə and *u, *o, *a, which does not exclude an original RTRbased contrast. Whereas Vovin (1993: 50-51) and Bugaeva (2015: 26-28) reconstruct palatal harmony in Ainu, Shibatani (1990: 15) speculates that the Ainu opposition between o and u, a might have its origin in tongue root harmony, but here the indications are even weaker than in the Japanese case.…”