“…It is involved in the composition of cells, the absorption and transport of fat‐soluble vitamins, energy supplement and the supply of essential fatty acids (Chen, Xu, et al., 2020; Chen, Sun, et al., 2020). Traditionally, fish oil is the most significant lipid source in the commercial feed of most aquatic animals because of its rich n−3 highly unsaturated fatty acid (HUFA), including eicosapentaenoic acid (EPA, C20:5n−3) and docosahexaenoic acid (DHA, C22:6n−3), which are essential fatty acids for cultured fish and crustaceans (Chen, Xu, et al., 2020; Chen, Sun, et al., 2020; Tocher, 2003, 2015). For C. quadricarinatus , plant oil can be a good substitute for fish oil because of the preference for phytophagous (Li et al., 2011; Thompson et al., 2010).…”