“…However, only a few peroxisome proteins are also implicated in chlorophyll biogenesis. For example, Peroxisomal Biogenesis Factor 2 (PEX2), a peroxisome protein involved in peroxisome assembly and matrix protein import, is required for photomorphogenesis in Arabidopsis [12]. The mutation of SNOWY COTYLEDON 3 (sco3), localized to the periphery of peroxisomes, resulted in reduced chlorophyll accumulation and defective chloroplast development, possibly through interaction with the cytoskeleton [13].…”