“…The earliest undisputed actinopterygians are from the middle to late Devonian (Janvier, 1996). Articulated remains of taxa such as Cheirolepis (Pearson & Westoll, 1979) and Moythomasia (Gross, 1950; Jessen, 1968, 1972; Gardiner, 1984) have palaeoniscoid scales (Fig. 12B,C), consisting of a superficial region of multilayered ganoine covering a region of dentine, and a thick basal plate of vascularized bone (Goodrich, 1907; Ørvig, 1978a,b,c; Sire, 1990).…”