Dedicated to Jim Cushing for all the inspiration, ever since Cortona 1979.We motivate and describe a class of nonlinear Leslie matrix models for semelparous populations, like cicadas and Pacific salmon. We then focus on a Cushing-inspired special case for which one can show rigorously that a heteroclinic boundary cycle exists and attracts nearby orbits for a certain range of parameters. Along the way we formulate some open problems concerning a carrying simplex and global behaviour.Keywords: Heteroclinic cycle; Leslie matrix model; Semelparous species
What is a year class?A species is called semelparous if reproduction also amounts to signing one's own death sentence. When there is exactly one reproduction opportunity per year (usually in the spring), we take one year as a natural discrete time unit. For several species, in particular many cicada species, the period in between being born and going to reproduce is strictly fixed at, say, k years. The population then subdivides into subpopulations according to the year of birth modulo k (or, equivalently, the year of reproduction modulo k). Such a subpopulation is called a year class (or, also, a brood). Note that whereas the age class to which an individual belongs increases by one when a year has passed, the year class to which it belongs is fixed once and for all.Year classes only mate among themselves and hence are reproductively isolated. In particular, if a year class goes extinct, it remains extinct. We then say that the year class is "missing". The periodical insects [1] are those for which all but one year classes are missing. The prime example is the Magicicadas (the k ¼ 17 species had its most recent emergence in the North-East United States in 2004).