“…According to Halffter (1991), Scarabaeinae is represented by a few abundant species and many species with few individuals in Neotropical forests. This pattern has been often found in studies conducted in forest ecosystems (e.g., Louzada and Lopes 1997, Vazde-Mello 1999, Medri and Lopes 2001, Milhomem et al 2003, Endres et al 2007, Louzada et al 2007, Filgueiras et al 2009, Hernández and Vaz-de-Mello 2009, Audino et al 2011, Lopes et al 2011, Silva 2011, in areas of fi elds or pastures in Brazil (e.g., Koller et al 2007, Matavelli and Louzada 2008, Louzada and Silva 2009, Lopes et al 2011, Silva 2011, as well as other Neotropical countries such as Bolivia (Spector and Ayzama 2003), Colombia (Martínez et al 2009), Ecuador (Peck and Forsyth 1982), Mexico (Halffter et al 1992) and Uruguay (Morelli et al 2002).…”