“…The second pathway, population density decline, is frequently reported (Tozer, Cameron, & Thom, 2011;Tozer et al, 2014), especially where environmental conditions are marginal or turn unfavourable for the sown species (Chapman, Edwards, & Nie, 2011). This pathway is the basis for measures such as ground cover scores to assess sward productivity (Camlin & Stewart, 1976) and to compare the persistence of grass cultivars (e.g., in perennial ryegrass; Cashman, O'Donovan, Gilliland, & McEvoy, 2014;O'Donovan, McHugh, McEvoy, Grogan, & Shalloo, 2016). An important implicit assumption is that the rate of perennial grass tiller mortality exceeds the rate of tiller replacement (dominantly through clonal reproduction and site filling rates, Davies, 1976) such that the population of perennial grass tillers cannot be sustained and herbage production declines (Camlin & Stewart, 1978).…”