“…E1 is phosphorylated by several kinases, including Cdk2, CK2, MAPK, and PKC [4, 12, 15, 19–22, 25, 33, 40, 41]. Three BPV E1 phosphorylation sites (serines 94, 95, and 100) are within the bipartite nuclear localization signal (NLS), and all three are phosphorylated by CK2 [22, 33].…”