“…Earlier comparative studies also indicated that these four genera, especially Nitraria and also Peganum (with Malacocarpus ), were more distinct from the remainder of Zygophyllaceae than previously suggested (for vegetative anatomy , see Sheahan and Chase, 1993 ; for palynology, see Erdtman, 1952 ;Agababyan, 1964 ;Perveen and Qaiser, 2006 ; for embryology, see Mauritzon, 1934 ;Sou è ges, 1953 ;Kapil and Ahluwalia, 1963 ;Kamelina, 1985Kamelina, , 1994Li and Tu, 1994 ; for fl oral structure, see Baillon, 1873 ;Nair and Nathawat, 1958 ;Ronse De Craene and Smets, 1991 ; ; for chemistry, see Hussein et al, 2009 ). In contrast to most of the current Zygophyllaceae, which have pentamerous, isomerous and obdiplostemonous fl owers, the fl owers in Tetradiclis are tetramerous, isomerous and haplostemonous , while in Nitraria and Peganum (+ Malacocarpus ), the fl owers have a pentamerous perianth and androecium, conspicuous antepetalous stamen pairs and a tricarpellate gynoecium (Payer, 1857 ;Baillon, 1873 ;Engler, 1896aEngler, , b , 1931Ronse De Craene and Smets, 1991 ;.…”