“…Since mimicry has obvious survival benefits, the above phenomenon raises the question of how nonmimetic individuals are maintained in populations without being completely replaced by mimetic individuals. This problem has been the focus of many researchers since Darwin's era (e.g., Bates, 1862; Komata, Lin, Iijima, Fujiwara, & Sota, 2016; Kunte, 2009b; Ohsaki, 1995, 2005; Ruxton et al, 2004; Turner, 1978; Wallace, 1867; Westerman et al., 2018, 2019; Wickler, 1968; Zhang, Westerman, Nitzany, Palmer, & Kronforst, 2017).…”