Beetles (Coleoptera) of Peru: A Survey of the Families. Scarabaeoidea

Abstract

Peruvian biodiversity is pivotal for conserving and managing natural resources, food security, poverty reduction, health, biosecurity, new industrial product development, and ecotourism (Smith et al., 2011). Geotrupidae Diversity in Peru: 1 subfamily, 4 genera, and 11 species. Recognition: The body shape is oval or round, and the head is not deflexed. The antennae are 11-segmented with a 3-segmented, opposable club with all antennomeres tomentose. The eyes are completely or partially divided by a canthus. The clypeus is often with a tubercle or horn. The labrum is truncate, prominent, and produced beyond the apex of the clypeus. The mandibles are prominent and produced beyond the apex of the labrum. The pronotum is convex with a base wider than or subequal to the elytral base and with or without tubercles, ridges, horns, or sulci. The elytra are convex, with or without striae. The pygidium is concealed by the elytra (Jameson, 2002a). Habitat: Life histories of the geotrupids are diverse, and food habits vary from saprophagous to coprophagous and mycetophagous. Adults of most species are secretive, living most of their life in burrows. Although adults do not tend larvae, adults provision food for larvae in brood burrows. Adults dig vertical burrows (15-200 cm in depth) and provision larval cells with dead leaves, cow dung, horse dung, or humus. Burrows of some species extend to a depth of 3.0 m (Jameson, 2002a). Notes: The family Geotrupidae includes 68 genera and about 620 species (Scholtz and Browne, 1996). The subfamily Geotrupinae does not occur in South America.

Synopsis of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae)

“…This striking, relatively large cyclocephaline occurs in lowland forests in Brazil, Ecuador, French Guiana, Peru, Suriname, and possibly Colombia (Burmeister 1847, Harold 1869b, Arrow 1937b, Blackwelder 1944, Endrődi 1966, 1985a, Endrődi and Dechambre 1976, Lachaume 1992, Couturier and Kahn 1992, Andreazze 2001, Andreazze and da Silva Motta 2002, Touroult et al 2010, Ponchel 2011, Saltin and Ratcliffe 2012, Ratcliffe et al 2015) (Fig. 60).…”

“…Ratcliffe has authored or coauthored 39 publications that cover cyclocephaline scarabs, and many of these are monographic in scope (Ratcliffe 1977, 1978, 1981, 1985, 1986, 1989, 1991, 1992a–d, 2002a, b, 2003, 2008, 2014, 2015, Ratcliffe and Cave 2002, 2006, 2008, 2009, 2010, 2015, 2017, Ratcliffe et al 2013, 2015, Ratcliffe and Delgado-Castillo 1990, Ratcliffe and Hoffman 2011, Ratcliffe and Morón 1997, Ratcliffe and Paulsen 2008, Figueroa and Ratcliffe 2016, Gasca-Álvarez et al 2014, Jameson et al 2002, 2009, Maes and Ratcliffe 1996, Maes et al 1997, Neita-Moreno et al 2006, 2007, Saltin and Ratcliffe 2012). …”

“…The five species and subspecies of Augoderia are distributed in Argentina, Bolivia, Brazil, French Guiana, Peru, and Venezuela (Burmeister 1847, Harold 1869b, Arrow 1937b, Blackwelder 1944, Guimarães 1944, Martínez 1966, Gibbs et al 1977, Endrődi 1966, 1967a, 1981, 1985a, Riehs 2005, Ronqui and Lopes 2006, Ponchel 2009, Grossi et al 2011, Ratcliffe et al 2015) (Fig. 55).…”

Actualización a la lista de coleópteros (Insecta: Coleoptera) de la región Piura, Perú

“…Las especies consideradas en esta actualización se presentan a modo de listado faunístico comentado, siendo la secuencia del material estudiado la siguiente: número de ♂♂, ♀♀, provincia, distrito, localidad, coordenadas geográficas, altitud en msnm, fecha de captura, tipo de captura, recolector, tipo de paisaje ecológico, código de registro cuando procedió de material depositado en la colección GJN y de una referencia para indicar los autores de los trabajos revisados cuando se recopiló a través de bibliografía. Las nuevas especies añadidas para la región se indican con una cruz ( †), mientras que las especies conocidas únicamente de Perú (Casari, 2002;González, 2015;Ratcliffe et al, 2015;Smith et al, 2015;Chandler, 2016;Giraldo & Flores, 2016;Pinto & Bologna, 2016;) se indican con un asterisco (*). Asimismo, se aporta una lista actualizada de todos los coleópteros conocidos hasta el momento para la región Piura incluyendo para cada especie su distribución por provincia y paisaje ecológico.…”

A new cryptic species and review of the east-Andean leaf chafer genus Mesomerodon Ohaus, 1905 (Coleoptera, Scarabaeidae, Rutelinae)

“…BOLIVIA: Cochabamba (Chapare), Huánuco (Blackwelder 1944, Ohaus 1905, 1918, 1934, 1952, Machatschke 1972, Soula 2008, Ratcliffe et al 2015), Junín (Blackwelder 1944, Ohaus 1905, 1918, 1934, 1952, Machatschke 1972, Soula 2008, Ratcliffe et al 2015), Pasco (Blackwelder 1944, Ohaus 1905, 1918, 1934, 1952, Machatschke 1972, Soula 2008, Ratcliffe et al 2015), Santa Cruz (Ichilo, Sara)…”