Magnetic reconnection is a key process in collisionless plasmas that converts magnetic energy into plasma kinetic and thermal energies through a rapid change of magnetic field topology. At the Earth's magnetopause, the reconnection process leads to efficient transport of solar wind energy into the magnetosphere, whereas in the magnetotail, it explosively releases the accumulated magnetic energy and causes global geomagnetic disturbances leading to auroral substorms (Baker et al., 1996;Hones, 1984). The energy conversion and topological changes take place in a very small region surrounding an X-line where ions or both ions and electrons are decoupled from the magnetic field. These regions are called the diffusion regions. They have a multiscale structure due to different masses and hence dynamics of ions and electrons. A larger ion diffusion region (IDR) with unmagnetized ions usually contains a smaller-scale electron diffusion region (EDR) where both electrons and ions are unmagnetized.A distinctive feature of the magnetotail reconnection onset is that corresponding plasma motions are needed to reduce the original finite normal magnetic field for the formation of an X-line. This is not necessary in case of antiparallel or sheared magnetic field configurations, such as at the magnetopause. Indeed, there is some controversy regarding whether fast plasma divergent flows in the magnetotail are a consequence or a cause of the magnetic topology change. In the former and the most commonly accepted scenario, the magnetic topology first changes because of the slow evolution of the tail induced by the external driving and subsequent microscale tearing instability (e.g., Liu et al., 2014). Therefore, plasma divergent flows arise from the unbalanced magnetic tension of the sharply kinked, newly reconnected field lines around the X-line.The latter scenario was originally proposed by Lin and Swift (2002). On the basis of the analysis of their 2-D global hybrid simulations, they conjectured that the magnetic topology change can be internally driven by the spontaneous generation of plasma divergent flows. The idea of such flows preceding and likely driving the reconnection onset was later elaborated on by Siscoe et al. (2009) andTanaka et al. (2019) in the analysis of their magnetohydrodynamic (MHD) simulations. These plasma divergent flows are referred to below as plasma "watersheds" (WSs), to distinguish them from reconnection ejecta emanated from an already formed X-line. Most recently, Sitnov, Motoba, and Swisdak (2021), using fully kinetic 3-D particle-in-cell (PIC) simulations,