“…1b), or intermediate (Fig. 1c; Arnold & Hodges, 1995; Coyne & Orr, 2004; Wellhausen, 1952; Edmands, 2007; Hatfield & Schluter, 1999; Fraïsse et al., 2016, Table ; Coughlan & Matute, 2020; Favre et al., 2017; Gramlich et al., 2022; Thompson & Schluter, 2022). Moreover, as illustrated in Figure 1d, the direction of the cross may be important, with strong fitness differences between the reciprocal F1 (i.e., female‐male vs. male‐female cross directions of the same parental lines).…”