“…Matelea sensu Woodson comprises at least 260 species, and several attempts have been made to describe smaller natural genera (e.g. Stevens 2001;Morillo 2012Morillo , 2013Morillo , 2015Morillo & Krings 2014). However, as Krings (2011: 730) stated, "The synorganization of androecium and gynoecium appears to have facilitated such a myriad of gynostegial structural variation … that it is at times difficult to discern relationships, much less morphological synapomorphies, in the absence of a resolved phylogeny".…”