“…Teeth we identify as similar to Dakotasuchus are conical, sometimes slightly lingually recurved, and typically sub-circular in occlusal view ( Figs. 7U – 7Y ) ( Frederickson et al, 2017 ). They are somewhat labiolingually compressed, with the labial side slightly more convex than the lingual.…”
Section: Systematic Paleontologymentioning
confidence: 99%
“…Enamel ornamentation is characterized by multiple, weakly defined, apicobasally oriented, parallel striations. Teeth of Dakotasuchus are morphologically similar to goniopholid and pholidosaurid teeth ( Garrison et al, 2007 ; Puértolas-Pascual, Rabal-Garcés & Canudo, 2015 ; Frederickson et al, 2017 ).…”
The vertebrate fauna of the Late Cretaceous Mussentuchit Member of the Cedar Mountain Formation has been studied for nearly three decades, yet the fossil-rich unit continues to produce new information about life in western North America approximately 97 million years ago. Here we report on the composition of the Cliffs of Insanity (COI) microvertebrate locality, a newly sampled site containing perhaps one of the densest concentrations of microvertebrate fossils yet discovered in the Mussentuchit Member. The COI locality preserves osteichthyan, lissamphibian, testudinatan, mesoeucrocodylian, dinosaurian, metatherian, and trace fossil remains and is among the most taxonomically rich microvertebrate localities in the Mussentuchit Member. To better refine taxonomic identifications of isolated theropod dinosaur teeth, we used quantitative analyses of taxonomically comprehensive databases of theropod tooth measurements, adding new data on theropod tooth morphodiversity in this poorly understood interval. We further provide the first descriptions of tyrannosauroid premaxillary teeth and document the earliest North American record of adocid remains, extending the appearance of this ancestrally Asian clade by 5 million years in western North America and supporting studies of pre-Cenomaninan Laurasian faunal exchange across Beringia. The overabundance of mesoeucrocodylian remains at the COI locality produces a comparatively low measure of relative biodiversity when compared to other microvertebrate sites in the Mussentuchit Member using both raw and subsampling methods. Much more microvertebrate research is necessary to understand the roles of changing ecology and taphonomy that may be linked to transgression of the Western Interior Seaway or microhabitat variation.
“…Teeth we identify as similar to Dakotasuchus are conical, sometimes slightly lingually recurved, and typically sub-circular in occlusal view ( Figs. 7U – 7Y ) ( Frederickson et al, 2017 ). They are somewhat labiolingually compressed, with the labial side slightly more convex than the lingual.…”
Section: Systematic Paleontologymentioning
confidence: 99%
“…Enamel ornamentation is characterized by multiple, weakly defined, apicobasally oriented, parallel striations. Teeth of Dakotasuchus are morphologically similar to goniopholid and pholidosaurid teeth ( Garrison et al, 2007 ; Puértolas-Pascual, Rabal-Garcés & Canudo, 2015 ; Frederickson et al, 2017 ).…”
The vertebrate fauna of the Late Cretaceous Mussentuchit Member of the Cedar Mountain Formation has been studied for nearly three decades, yet the fossil-rich unit continues to produce new information about life in western North America approximately 97 million years ago. Here we report on the composition of the Cliffs of Insanity (COI) microvertebrate locality, a newly sampled site containing perhaps one of the densest concentrations of microvertebrate fossils yet discovered in the Mussentuchit Member. The COI locality preserves osteichthyan, lissamphibian, testudinatan, mesoeucrocodylian, dinosaurian, metatherian, and trace fossil remains and is among the most taxonomically rich microvertebrate localities in the Mussentuchit Member. To better refine taxonomic identifications of isolated theropod dinosaur teeth, we used quantitative analyses of taxonomically comprehensive databases of theropod tooth measurements, adding new data on theropod tooth morphodiversity in this poorly understood interval. We further provide the first descriptions of tyrannosauroid premaxillary teeth and document the earliest North American record of adocid remains, extending the appearance of this ancestrally Asian clade by 5 million years in western North America and supporting studies of pre-Cenomaninan Laurasian faunal exchange across Beringia. The overabundance of mesoeucrocodylian remains at the COI locality produces a comparatively low measure of relative biodiversity when compared to other microvertebrate sites in the Mussentuchit Member using both raw and subsampling methods. Much more microvertebrate research is necessary to understand the roles of changing ecology and taphonomy that may be linked to transgression of the Western Interior Seaway or microhabitat variation.
“…One possible explanation for this distribution is that these medium-sized theropods were competitively excluded from the near-water settings by large-bodied coelognathosuchian crocodilians. Some of these species potentially reached sizes over 5 m 20 , and their teeth are some of the most commonly encountered vertebrate fossils found in the MM 19 . Alternative hypotheses ranging from poor pedal traction near water to overgrown hunting terrain could also be viable explanations for this distribution.…”
Section: Discussionmentioning
confidence: 99%
“…The MM formed as shed sediments deposited in the foreland basin during the Pavast thrust event, dated from 98.3 ± 0.1 Ma 18 to 96.7 ± 0.5 Ma 19 , placing the member in the Cenomanian stage of the Late Cretaceous. The unit is generally highly fossiliferous (as compared to the rest of the Cedar Mountain Formation), with a diverse fauna composed of freshwater fishes, lissamphibians, lepidosaurs, crocodilians, dinosaurs, and mammals 19,20 .Figure 1Map of the western interior during the Cenomanian (grey is highlands, green is lowlands, and blue is water) and a pullout of Emery County, Utah with the Mussentuchit Member exposure and microsites analyzed in this study. Based on maps and data from Cifelli et al .…”
We explore hypothetical ecologies to explain diversity among predatory dinosaurs in North America’s medial Cretaceous, based on occurrence, tooth morphology, and stable isotope analysis. The Mussentuchit local fauna, Utah, USA, is among the best-known terrestrial vertebrate assemblages from the Cretaceous. Study samples include teeth from six microvertebrate sites, ranging in depositional setting from distal floodplain to channel lags. We recognize four theropod morphotypes: a comparatively large theropod (morph 1), a medium-sized dromaeosaurid (morph 2), a small dromaeosaurid (morph 3), and a tooth-morph similar to the genus Richardoestesia (morph 4). These four morphotypes vary significantly in mean size, from 15.1 mm in the largest theropod to 3.7 mm in Richardoestesia. Further, tooth representation from two of the best-sampled microsites (representing a channel/splay and floodplain deposit) show differing patterns of abundances with morphs 1 and 3 having roughly the same abundance in both sites, while morph 2 was more abundant in the floodplain setting and morph 4 was more abundant in the channel/splay. Stable isotope analysis (δ13C; δ18O) of tooth carbonate from the theropod morphotypes, goniopholidid crocodilians, and matrix (to test for diagenesis) from these sites were also analyzed. The theropods show modest differences in δ13C values between each other, with carbonate from the teeth of morphs 1, 3, and 4 being enriched in 13C for the channel/splay relative to the floodplain environments, possibly indicative of dietary plasticity in these species. We hypothesize that these data indicate that the Mussentuchit theropods had different niches within the predator guild, suggesting plausible means by which ecospace was divided among the predatory dinosaurs of the Mussentuchit local fauna.
The Crocodyliformes are the most represented vertebrate clade in the Upper Cretaceous sequences of the Bauru Group, Paraná Basin. However, some of the species described have an uncertain taxonomic status and phylogenetic position. For instance, “Goniopholis” paulistanus has been assigned as a nomem dubium, due to its description being based on scarce material. The “G”. paulistanus specimens (i.e. teeth and a left tibia) were discovered in two different localities in São Paulo state: Mirandópolis and Valparaíso municipalities; where the upper interval of the Adamantina Formation (Early Maastrichtian of Bauru Group) crops out. Revisiting these specimens, we observed multicrenulated teeth in middle dentary toot- row, a remarkable feature only shared with teleosaurids Machimosaurus hugii (Upper Jurassic of Laurasia) and M. rex (Lower Cretaceous of Tunisia). This apomorphy was also recognized in new material from the Alfredo Marcondes municipality (Presidente Prudente Formation), which are here also referred to “G”. paulistanus. We recognized the teeth of “G.” paulistanus as the lectotype, however the tibia cannot be assigned to a species as it was not collected in association with the teeth. We performed a phylogenetic analysis with a data matrix composed of 388 characters and 86 taxa, analyzed in TNT. The strict consensus tree recovered Neosuchia and Ziphosuchia (Notosuchia + Sebecia) within Mesoeucrocodylia. The species “G” paulistanus is valid, as a distinct and new genus within Sebecia, in a polytomy with Barreirosuchus, Pepesuchus, Itasuchus and Peirosaurus, forming the clade Itasuchidae. Stolokrosuchus is the sister taxon to Itasuchidae, the sister group of all other Sebecia (Peirosauridae (Mahajangasuchidae + Sebecidae and taxa affinis)). The clades Ziphosuchia, Sebecia and Itasuchidae are here redefined, and we find the last two clades to be more closely related to terrestrial notosuchids than to semiaquatic neosuchians.
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