“…Tristes Singer and successively placed by Smith and Thiers [23] in Tylopilus sect. Graciles A.H. Smith and Thiers. The genus as presently outlined is characterized from the morphological viewpoint by boletoid fruiting bodies with dry to viscid or even mucilaginous pileus and stipe surfaces, initially whitish or pale cream becoming flesh-pink to vinaceous pink or brownish pink tubular hymenophore at maturity, smooth, furfuraceous-fibrillose to more often markedly reticulate-alveolate, lacerate or lacunose stipe, generally unchanging tissues, flesh-pink, pinkish vinaceous, purplish brown, rust brown to chocolate brown spore print, variously ornamented (finely verrucose or warted to irregularly pitted but also flat-tuberculate to subreticulate) amygdaliform to ellipsoid-fusiform basidiospores, trichoderm or ixotrichoderm pileipellis, bilateral-divergent hymenophoral trama of the "Boletus-type", gymnocarpic, velangiocarpic (primary angiocarpy), or pseudoangiocarpic (secondary angiocarpic) ontogenesis and ectomycorrhizal (ECM) association with several plant families including Fagaceae, Pinaceae, Dipterocarpaceae, Myrtaceae, and caesalpinoid legumes [8,9,12,14,[16][17][18]20,21,[24][25][26][27][28][29][30][31][32][33][34][35], although some species are suspected to be saprotrophic or only facultative ECM [32]. Austroboletus appears to be scarcely represented in temperate woodlands of both hemispheres but is particularly diverse throughout the pantropical belt, especially across the neotropical latitudes of Central and northern South America and all along the Australasian region [9,16,18,20,26,30,[32][33][34]36...…”