“…Some authors consider them to be two different species (Tostain et al , 2007; Bressan et al , 2014), while others consider both as part of a species complex (Dansi et al , 2001, 2013; Mignouna et al , 2002; Zannou et al , 2009; Girma et al , 2014). Several studies have used morphological markers to measure the genetic diversity of these species (Onyilagha, 1986; Dansi et al , 1999; Melo Filho et al , 2000; Bressan et al , 2014; Girma et al , 2014; Loko et al , 2015). Among the molecular markers, microsatellites or SSR (simple sequence repeats) stand out for their efficiency, and for being multi-allelic, co-dominant, and showing a simple Mendelian segregation (Agarwal et al , 2008).…”