“…More specifically, Mohr (1943) described dung pads as scattered microhabitats of high-quality resources of relatively small size and short duration that support complex communities. Factors that collectively influence the composition of dung beetle communities include soil and vegetation type (Lumaret and Kirk, 1987;Krell et al, 2003), source of dung (herbivore, carnivore, and omnivore) (Dormont et al, 2007;Filgueiras et al, 2009;Whipple, 2011), age of dung (Barth et al, 1994;Kryger et al, 2006), size of pad (Peck and Howden, 1984;Finn and Giller, 2000), soil moisture/rainfall events (Kingston, 1977;Lumaret, 1978), altitude (Mittal, 1981;Avila and Pascual, 1987), latitude (Kirk and Ridsdill-Smith, 1986;Hanski, 1991), mean temperature/season (Lumaret, 1979;Davis, 1989), and diel activity (Hernandez, 2002;Krell et al, 2003).…”