Climatic variability is a dominant factor affecting large wildfires in the western United States, an observation supported by palaeoecological data on charcoal in lake sediments and reconstructions from fire-scarred trees. Although current fire management focuses on fuel reductions to bring fuel loadings back to their historical ranges, at the regional scale extreme fire weather is still the dominant influence on area burned and fire severity. Current forecasting tools are limited to short-term predictions of fire weather, but increased understanding of large-scale oceanic and atmospheric patterns in the Pacific Ocean (e.g., El Niño Southern Oscillation, Pacific Decadal Oscillation) may improve our ability to predict climatic variability at seasonal to annual leads. Associations between these quasi-periodic patterns and fire occurrence, though evident in some regions, have been difficult to establish in others. Increased temperature in the future will likely extend fire seasons throughout the western United States, with more fires occurring earlier and later than is currently typical, and will increase the total area burned in some regions. If climatic change increases the amplitude and duration of extreme fire weather, we can expect significant changes in the distribution and abundance of dominant plant species in some ecosystems, which would thus affect habitat of some sensitive plant and animal species. Some species that are sensitive to fire may decline, whereas the distribution and abundance of species favored by fire may be enhanced. The effects of climatic change will partially depend on the extent to which resource management modifies vegetation structure and fuels. Resumen: La variabilidad climática es un factor dominante que afecta a incendios mayores en el oestede Estados Unidos, observación sustentada por datos paleoecológicos de carbón en sedimentos lacustres y reconstrucciones deárboles con cicatrices de fuego. Aunque la gestión actual de incendios se centra en la reducción de combustibles para que las cargas de combustible retornen a sus valores históricos, en la escala regional el climaígneo extremo aun es la influencia dominante delárea quemada e intensidad del fuego. Las herramientas predictivas actuales están limitadas a pronósticos de climaígneo a corto plazo, pero mejorar nuestra habilidad para predecir la variabilidad climática desde estacional a anualmente podría mejorar con mayor conocimiento de los patrones oceánicos y atmosféricos a gran escala en el Océano Pacífico (por ejemplo, Oscilación Meridional El Niño, Oscilación Decadal del Pacífico). La asociación entre estos patrones cuasi-periódicos y la ocurrencia de incendios, aunque evidente en algunas regiones, ha sido difícil de establecer en otras. El futuro incremento de temperatura probablemente extenderá las temporadas de incendios en el oeste de Estados Unidos, con más incendios antes y después de lo típico actualmente, e incremento en la superficie quemada en algunas regiones. Si el cambio climático incrementa la amplitud y duraci...

Before the advent of intensive forest management and fire suppression, western North American forests exhibited a naturally occurring resistance and resilience to wildfires and other disturbances. Resilience, which encompasses resistance, reflects the amount of disruption an ecosystem can withstand before its structure or organization qualitatively shift to a different basin of attraction. In fire-maintained forests, resilience to disturbance events arose primarily from vegetation pattern-disturbance process interactions at several levels of organization. Using evidence from 15 ecoregions, spanning forests from Canada to Mexico, we review the properties of forests that reinforced qualities of resilience and resistance. We show examples of multi-level landscape resilience, of feedbacks within and among levels, and how conditions have changed under climatic and management influences. We highlight geographic similarities and important differences in the structure and organization of historical landscapes, their forest types, and in the conditions that have changed resilience and resistance to abrupt or large-scale Hessburg et al. Resilience in North American Forests disruptions. We discuss the role of the regional climate in episodically or abruptly reorganizing plant and animal biogeography and forest resilience and resistance to disturbances. We give clear examples of these changes and suggest that managing for resilient forests is a construct that strongly depends on scale and human social values. It involves human communities actively working with the ecosystems they depend on, and the processes that shape them, to adapt landscapes, species, and human communities to climate change while maintaining core ecosystem processes and services. Finally, it compels us to embrace management approaches that incorporate ongoing disturbances and anticipated effects of climatic changes, and to support dynamically shifting patchworks of forest and non-forest. Doing so could make these shifting forest conditions and wildfire regimes less disruptive to individuals and society.

Wildland fire is an important disturbance agent in forests of the American Northwest. Historical fire suppression efforts have contributed to an accumulation of fuels in many Northwestern forests and may result in more frequent and/or more severe wildfire events. Here we investigate the extent to which atmospheric and climatic variability may contribute to variability in annual area burned on 20 National Forests in Washington, Oregon, and Idaho. Empirical orthogonal function (EOF) analysis was used to identify coherent patterns in area burned by wildfire in the Pacific Northwest. Anomaly fields of 500‐hPa height were regressed onto the resulting principal‐component time series to identify the patterns in atmospheric circulation that are associated with variability in area burned by wildfire. Additionally, cross‐correlation functions were calculated for the Palmer drought severity index (PDSI) over the year preceding the wildfire season. Parallel analyses based on superposed epoch analysis focused only on the extreme fire years (both large and small) to discriminate the controls on extreme years from the linear responses identified in the regression analyses. Four distinct patterns in area burned were identified, each associated with distinct climatic processes. Extreme wildfire years are forced at least in part by antecedent drought and summertime blocking in the 500‐hPa height field. However the response to these forcings is modulated by the ecology of the dominant forest. In more mesic forest types antecedent drought is a necessary precondition for forests to burn, but it is not a good predictor of area burned due to the rarity of subsequent ignition. At especially dry locations, summertime blocking events can lead to increases in area burned even in the absence of antecedent drought. At particularly xeric locations summertime cyclones can also lead to increased area burned, probably due to dry lightning storms that bring ignition and strong winds but little precipitation. These results suggest that fuels treatments alone may not be effective at reducing area burned under extreme climatic conditions and furthermore that anthropogenic climate change may have important implications for forest management.

Though tree-ring chronologies are annually resolved, their dating has never been independently validated at the global scale. Moreover, it is unknown if atmospheric radiocarbon enrichment events of cosmogenic origin leave spatiotemporally consistent fingerprints. Here we measure the 14C content in 484 individual tree rings formed in the periods 770–780 and 990–1000 CE. Distinct 14C excursions starting in the boreal summer of 774 and the boreal spring of 993 ensure the precise dating of 44 tree-ring records from five continents. We also identify a meridional decline of 11-year mean atmospheric radiocarbon concentrations across both hemispheres. Corroborated by historical eye-witness accounts of red auroras, our results suggest a global exposure to strong solar proton radiation. To improve understanding of the return frequency and intensity of past cosmic events, which is particularly important for assessing the potential threat of space weather on our society, further annually resolved 14C measurements are needed.

Temperature is a primary driver of the distribution of biodiversity as well as of ecosystem boundaries. Declining temperature with increasing elevation in montane systems has long been recognized as a major factor shaping plant community biodiversity, metabolic processes, and ecosystem dynamics. Elevational gradients, as thermoclines, also enable prediction of long-term ecological responses to climate warming. One of the most striking manifestations of increasing elevation is the abrupt transitions from forest to treeless alpine tundra. However, whether there are globally consistent above- and belowground responses to these transitions remains an open question. To disentangle the direct and indirect effects of temperature on ecosystem properties, here we evaluate replicate treeline ecotones in seven temperate regions of the world. We find that declining temperatures with increasing elevation did not affect tree leaf nutrient concentrations, but did reduce ground-layer community-weighted plant nitrogen, leading to the strong stoichiometric convergence of ground-layer plant community nitrogen to phosphorus ratios across all regions. Further, elevation-driven changes in plant nutrients were associated with changes in soil organic matter content and quality (carbon to nitrogen ratios) and microbial properties. Combined, our identification of direct and indirect temperature controls over plant communities and soil properties in seven contrasting regions suggests that future warming may disrupt the functional properties of montane ecosystems, particularly where plant community reorganization outpaces treeline advance.