12Human visual object recognition is subserved by a multitude of cortical areas. To make sense 13 of this system, one line of research focused on response properties of primary visual cortex 14 neurons and developed theoretical models of a set of canonical computations such as convolution, 15thresholding, exponentiating and normalization that could be hierarchically repeated to give 16 rise to more complex representations. Another line or research focused on response properties 17 of high-level visual cortex and linked these to semantic categories useful for object recognition. (which was not peer-reviewed) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity.