The spatio-temporal pattern of peak Holocene warmth (Holocene thermal maximum, HTM) is traced over 140 sites across the Western Hemisphere of the Arctic (0-180 W; north of B60 N). Paleoclimate inferences based on a wide variety of proxy indicators provide clear evidence for warmer-than-present conditions at 120 of these sites. At the 16 terrestrial sites where quantitative estimates have been obtained, local HTM temperatures (primarily summer estimates) were on average 1.670.8 C higher than present (approximate average of the 20th century), but the warming was time-transgressive across the western Arctic. As the precession-driven summer insolation anomaly peaked 12-10 ka (thousands of calendar years ago), warming was concentrated in northwest North America, while cool conditions lingered in the northeast. Alaska and northwest Canada experienced the HTM between ca 11 and 9 ka, about 4000 yr prior to the HTM in northeast Canada. The delayed warming in Quebec and Labrador was linked to the residual Laurentide Ice Sheet, which chilled the region through its impact on surface energy balance and ocean circulation. The lingering ice also attests to the inherent asymmetry of atmospheric and oceanic circulation that predisposes the region to glaciation and modulates the pattern of climatic change. The spatial asymmetry of warming during the HTM resembles the pattern of warming observed in the Arctic over the last several decades. Although the two warmings are described at different temporal scales, and the HTM was additionally affected by the residual Laurentide ice, the similarities suggest there might be a preferred mode of variability in the atmospheric circulation that generates a recurrent pattern of warming under positive radiative forcing. Unlike the HTM, however, future warming will not be counterbalanced by the cooling effect of a residual North American ice sheet. r ARTICLE IN PRESS

Knowledge of historical fire activity tends to be focused at local to landscape scales with few attempts to examine how local patterns of fire activity scale to global patterns. Generally, fire activity varied globally and continuously since the last glacial maximum (LGM) in response to long-term changes in global climate and shorter-term regional changes in climate, vegetation, and human land use. We have synthesised sedimentary charcoal records of biomass burning since the LGM and present global maps showing changes in fire activity for time slices during the past 21,000 years (as differences in charcoal accumulation values compared to pre-industrial). There is strong broad-scale coherence in fire activity after the LGM, but spatial heterogeneity in the signals increases thereafter. In eastern and western North America and western Europe and southern South America, charcoal records indicate less-than-present fire activity from 21,000 to ~11,000 cal yr BP. In contrast, the tropical latitudes of South America and Africa show greaterthan-present fire activity from ~19,000 to ~17,000 cal yr BP whereas most sites from Indochina and Australia show greater-than-present fire activity from 16,000 to ~13,000 cal yr BP. Many sites indicate greater-than-present or near-present activity during the Holocene with the exception of eastern North America and eastern Asia from 8000 to ~2000 cal yr BP, Indonesia from 11,000 to 4000 cal yr BP, and southern South America from 6000 to 3000 cal yr BP where fire activity was less than present. Regional coherence in the patterns of change in fire activity was evident throughout the postglacial period. These complex patterns can be explained in terms of large-scale climate controls modulated by local changes in vegetation and fuel load.

Reconstructing climates of the past relies on a variety of evidence from a large number of sites to capture the varied features of climate and the spatial heterogeneity of climate 32 change. This review summarizes available information from diverse Holocene paleoenvironmental records across eastern Beringia (Alaska, westernmost Canada and adjacent 34 seas), and it quantifies the primary trends of temperature-and moisture-sensitive records based in part on midges, pollen, and biogeochemical indicators (compiled in the recently published Arctic 36Holocene database, and updated here to v2.1). The composite time series from these proxy records are compared with new summaries of mountain-glacier and lake-level fluctuations, 38 terrestrial water-isotope records, sea-ice and sea-surface-temperature analyses, and peatland and thaw-lake initiation frequencies to clarify multi-centennial-to millennial-scale trends in 40Holocene climate change. To focus the synthesis, the paleo data are used to frame specific questions that can be addressed with simulations by Earth system models to investigate the 42 causes and dynamics of past and future climate change. This systematic review shows that, during the early Holocene (11.7-8.2 ka), rather than a prominent thermal maximum as suggested 44 previously, temperatures were highly variable, at times both higher and lower than present (approximate mid-20 th -century average), with no clear spatial pattern. Composited pollen, midge 46 and other proxy records average out the variability and show the overall lowest summer and mean-annual temperatures across the study region during the earliest Holocene, followed by 48 warming over the early Holocene. The sparse data available on early Holocene glaciation show that glaciers in southern Alaska were as extensive then as they were during the late Holocene. 50Early Holocene lake levels were low in interior Alaska, but moisture indicators show pronounced differences across the region. The highest frequency of both peatland and thaw-lake initiation 52 ages also occurred during the early Holocene. During the middle Holocene (8.2-4.2 ka), glaciers 3 retreated as the regional average temperature increased to a maximum between 7 and 5 ka, as 54 reflected in most proxy types. Following the middle Holocene thermal maximum, temperatures decreased starting between 4 and 3 ka, signaling the onset of Neoglacial cooling. Glaciers in the 56 Brooks and Alaska Ranges advanced to their maximum Holocene extent as lakes generally rose to modern levels. Temperature differences for averaged 500-year time steps typically ranged by 58 1-2°C for individual records in the Arctic Holocene database, with a transition to a cooler late Holocene that was neither abrupt nor spatially coherent. The longest and highest-resolution 60 terrestrial water isotope records previously interpreted to represent changes in the Aleutian lowpressure system around this time are here shown to be largely contradictory. Furthermore, there 62 are too few records with sufficient resolution to ...

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Abstract. The mid-Holocene decline of eastern hemlock is widely viewed as the sole prehistorical example of an insect-or pathogen-mediated collapse of a North American tree species and has been extensively studied for insights into pest-host dynamics and the consequences to terrestrial and aquatic ecosystems of dominant-species removal. We report paleoecological evidence implicating climate as a major driver of this episode. Data drawn from sites across a gradient in hemlock abundance from dominant to absent demonstrate: a synchronous, dramatic decline in a contrasting taxon (oak); changes in lake sediments and aquatic taxa indicating low water levels; and one or more intervals of intense drought at regional to continental scales. These results, which accord well with emerging climate reconstructions, challenge the interpretation of a biotically driven hemlock decline and highlight the potential for climate change to generate major, abrupt dynamics in forest ecosystems.