Logistic regression models used to predict tree mortality are critical to post-fire management, planning prescribed burns and understanding disturbance ecology. We review literature concerning post-fire mortality prediction using logistic regression models for coniferous tree species in the western USA. We include synthesis and review of: methods to develop, evaluate and interpret logistic regression models; explanatory variables in logistic regression models; factors influencing scope of inference and model limitations; model validation; and management applications. Logistic regression is currently the most widely used and available technique for predicting post-fire tree mortality. Over 100 logistic regression models have been developed to predict post-fire tree mortality for 19 coniferous species following wild and prescribed fires. The most widely used explanatory variables in post-fire tree mortality logistic regression models have been measurements of crown (e.g. crown scorch) and stem (e.g. bole char) injury. Prediction of post-fire tree mortality improves when crown and stem variables are used collectively. Logistic regression models that predict post-fire tree mortality are the basis of simple field tools and contribute to larger fire-effects models. Future post-fire tree mortality prediction models should include consistent definition of model variables, model validation and direct incorporation of physiological responses that link to process modelling efforts.
Restoring forest ecosystems has become an increasingly high priority for land managers across the American West. Millions of hectares of forest are in need of drastic yet strategic reductions in density (e.g., basal area). Meeting the restoration and management goals requires quantifying metrics of vertical and horizontal forest structure, which has relied upon field‐based measurements, manned airborne or satellite remote sensing datasets. We used unmanned aerial vehicle (UAV) image‐derived Structure‐from‐Motion (SfM) models and high‐resolution multispectral orthoimagery in this study to quantify vertical and horizontal forest structure at both the fine‐ (<4 ha) and mid‐scales (4–400 ha) across a forest density gradient. We then used these forest structure estimates to assess specific objectives of a forest restoration treatment. At the fine‐scale, we found that estimates of individual tree height and canopy diameter were most accurate in low‐density conditions, with accuracies degrading significantly in high‐density conditions. Mid‐scale estimates of canopy cover and forest density followed a similar pattern across the density gradient, demonstrating the effectiveness of UAV image‐derived estimates in low‐ to medium‐density conditions as well as the challenges associated with high‐density conditions. We found that post‐treatment conditions met a majority of the prescription objectives and demonstrate the UAV image application in quantifying changes from a mechanical thinning treatment. We provide a novel approach to forest restoration monitoring using UAV‐derived data, one that considers varying density conditions and spatial scales. Future research should consider a more spatially extensive sampling design, including different restoration treatments, as well as experimenting with different combinations of equipment, flight parameters, and data processing workflows.
Recent large scale mountain pine beetle (Dendroctonus ponderosae Hopkins, MPB) outbreaks have created concern regarding increased fuel loadings and exacerbated fire behavior and have prompted a desire to understand the effects of sequential disturbances on the landscape. However, previous research has focused on quantifying fuel loadings and using operational fire behavior models, rather than direct field measurements, to understand changes in fire severity following MPB. The 2012 Pole Creek Fire in central Oregon partially occurred in gray stage (8-15 years post-MPB epidemic) lodgepole pine forests. We examined the combined effects of MPB and fire disturbances on stand structure, and investigated the influence of previous MPB severity and fire weather on subsequent fire severity and cumulative disturbance severity. We randomly selected and installed 52 plots over a gradient of MPB and fire severity combinations and measured stand structure and fire severity characteristics. Fire severity metrics representing both crown and surface fire decreased with increased MPB severity under extreme burning conditions, following expected trends for crown fire severity, but not surface fire severity. Cumulative basal area mortality increased with MPB severity under moderate burning conditions, while other cumulative disturbance severity metrics were unrelated or weakly related to MPB severity. High severity crown fire was common despite hypothesized low canopy fuel loadings during the gray stage, indicating the importance of understanding variable mortality density of MPB outbreaks. Although long-term studies are needed to understand ecosystem recovery trajectories over time, there was no indication that a loss of ecosystem resilience occurred as a result of two sequential disturbances in this landscape.
Increasing fire severity and warmer, drier postfire conditions are making forests in the western United States (West) vulnerable to ecological transformation. Yet, the relative importance of and interactions between these drivers of forest change remain unresolved, particularly over upcoming decades. Here, we assess how the interactive impacts of changing climate and wildfire activity influenced conifer regeneration after 334 wildfires, using a dataset of postfire conifer regeneration from 10,230 field plots. Our findings highlight declining regeneration capacity across the West over the past four decades for the eight dominant conifer species studied. Postfire regeneration is sensitive to high-severity fire, which limits seed availability, and postfire climate, which influences seedling establishment. In the near-term, projected differences in recruitment probability between low- and high-severity fire scenarios were larger than projected climate change impacts for most species, suggesting that reductions in fire severity, and resultant impacts on seed availability, could partially offset expected climate-driven declines in postfire regeneration. Across 40 to 42% of the study area, we project postfire conifer regeneration to be likely following low-severity but not high-severity fire under future climate scenarios (2031 to 2050). However, increasingly warm, dry climate conditions are projected to eventually outweigh the influence of fire severity and seed availability. The percent of the study area considered unlikely to experience conifer regeneration, regardless of fire severity, increased from 5% in 1981 to 2000 to 26 to 31% by mid-century, highlighting a limited time window over which management actions that reduce fire severity may effectively support postfire conifer regeneration.
Higher tree density, more fuels, and a warmer, drier climate have caused an increase in the frequency, size, and severity of wildfires in western U.S. forests. There is an urgent need to restore forests across the western United States. To address this need, the U.S. Forest Service began the Four Forest Restoration Initiative (4FRI) to restore four national forests in Arizona. The objective of this study was to evaluate how restoration of ~400,000 ha under the 4FRI program and projected climate change would influence carbon dynamics and wildfire severity from 2010 to 2099. Specifically, we estimated forest carbon fluxes, carbon pools and wildfire severity under a moderate and fast 4FRI implementation schedule and compared those to status quo and no‐harvest scenarios using the LANDIS‐II simulation model and climate change projections. We found that the fast‐4FRI scenario showed early decreases in ecosystem carbon due to initial thinning/prescribed fire treatments, but total ecosystem carbon increased by 9–18% over no harvest by the end of the simulation. This increased carbon storage by 6.3–12.7 million metric tons, depending on the climate model, equating to removal of carbon emissions from 55,000 to 110,000 passenger vehicles per year until the end of the century. Nearly half of the additional carbon was stored in more stable soil pools. However, climate models with the largest predicted temperature increases showed declines by late century in ecosystem carbon despite restoration. Our study uses data from a real‐world, large‐scale restoration project and indicates that restoration is likely to stabilize carbon and the benefits are greater when the pace of restoration is faster.
1. The future of dry forests around the world is uncertain given predictions that rising temperatures and enhanced aridity will increase drought-induced tree mortality. Using forest management and ecological restoration to reduce density and competition for water offers one of the few pathways that forests managers can potentially minimize drought-induced tree mortality. Competition for water during drought leads to elevated tree mortality in dense stands, although the influence of density on heat-induced stress and the durations of hot or dry conditions that most impact mortality remain unclear.2. Understanding how competition interacts with hot-drought stress is essential to recognize how, where and how much reducing density can help sustain dry forests in a rapidly changing world. Here, we integrated repeat measurements of 28,881 ponderosa pine trees across the western US (2000)(2001)(2002)(2003)(2004)(2005)(2006)(2007)(2008)(2009)(2010)(2011)(2012)(2013)(2014)(2015)(2016)(2017) with soil moisture estimates from a water balance model to examine how annual mortality responds to competition, temperature and soil moisture conditions.3. Tree mortality responded most strongly to basal area, and was elevated in places with high mean temperatures, unusually hot 7-year high temperature anomalies, and unusually dry 8-year low soil moisture anomalies. Mortality was also lower in places that experienced unusually wet 3-year soil moisture anomalies between measurements. Importantly, we found that basal area interacts with temperature and soil moisture, exacerbating mortality during times of stress imposed by high temperature or low moisture.4. Synthesis and applications. Our results imply that a 50% reduction in forest basal area could reduce drought-driven tree mortality by 20%-80%. The largest impacts of density reduction are seen in areas with high current basal area and places that experience high temperatures and/or severe multiyear droughts. These interactions between competition and drought are critical to understand past and future patterns of tree mortality in the context of climate change, and provide information for resource managers seeking to enhance dry forest drought resistance.
Lodgepole pine (Pinus contorta) forests are widely distributed throughout North America and are subject to mountain pine beetle (Dendroctonus ponderosae) epidemics, which have caused mortality over millions of hectares of mature trees in recent decades. Mountain pine beetle is known to influence stand structure, and has the ability to impact many forest processes. Dwarf mistletoe (Arceuthobium americanum) also influences stand structure and occurs frequently in post-mountain pine beetle epidemic lodgepole pine forests. Few studies have incorporated both disturbances simultaneously although they co-occur frequently on the landscape. The aim of this study is to investigate the stand structure of lodgepole pine forests 21–28 years after a mountain pine beetle epidemic with varying levels of dwarf mistletoe infection in the Deschutes National Forest in central Oregon. We compared stand density, stand basal area, canopy volume, proportion of the stand in dominant/codominant, intermediate, and suppressed cohorts, average height and average diameter of each cohort, across the range of dwarf mistletoe ratings to address differences in stand structure. We found strong evidence of a decrease in canopy volume, suppressed cohort height, and dominant/codominant cohort diameter with increasing stand-level dwarf mistletoe rating. There was strong evidence that as dwarf mistletoe rating increases, proportion of the stand in the dominant/codominant cohort decreases while proportion of the stand in the suppressed cohort increases. Structural differences associated with variable dwarf mistletoe severity create heterogeneity in this forest type and may have a significant influence on stand productivity and the resistance and resilience of these stands to future biotic and abiotic disturbances. Our findings show that it is imperative to incorporate dwarf mistletoe when studying stand productivity and ecosystem recovery processes in lodgepole pine forests because of its potential to influence stand structure.
Wildland fires have a multitude of ecological effects in forests, woodlands, and savannas across the globe. A major focus of past research has been on tree mortality from fire, as trees provide a vast range of biological services. We assembled a database of individual-tree records from prescribed fires and wildfires in the United States. The Fire and Tree Mortality (FTM) database includes records from 164,293 individual trees with records of fire injury (crown scorch, bole char, etc.), tree diameter, and either mortality or top-kill up to ten years post-fire. Data span 142 species and 62 genera, from 409 fires occurring from 1981-2016. Additional variables such as insect attack are included when available. The FTM database can be used to evaluate individual fire-caused mortality models for pre-fire planning and post-fire decision support, to develop improved models, and to explore general patterns of individual fire-induced tree death. The database can also be used to identify knowledge gaps that could be addressed in future research.
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