The changing nature of the communities of boring and encrusting taxa found on upward‐facing hard‐grounds has been studied from the standpoints of (a) diversity, (b) faunal composition, and (c) nature of the niches occupied. After a rapid initial increase in the early Palaeozoic, diversity remained at much the same level from the Middle Ordovician until the late Cretaceous. However, there is a considerable turnover in the identity of the individual taxa between successive sample intervals. The incoming and outgoing of the major groups parallel their fortunes in the marine realm as a whole. Niche analysis suggests that the same feeding levels are occupied for most of the history of hardground communities, but Mesozoic faunas contain a much higher proportion of species with true exoskcletons, or which lived infaunally. The evolution of these forms was probably influenced by the Mesozoic radiation of marine predators and duriphages, but it also resulted in Mesozoic hardground faunas being more resistant than their Palaeozoic counterparts to episodic corrasion. Resulting higher population densities in the Mesozoic were probably one reason why cavity faunas beneath some of these hardground surfaces are more diverse than those beneath Palaeozoic examples. □Hardground, community, evolution.
Tim 1984 10 15: Commissural asymmetry in brachiopods. Lethaia. Vol. 17, Consistent asymmetric folding of the commissure is a characteristic feature of a small but significant number of brachiopod species. The feature may be obligate or facultative and is almost entirely confined to rhynchonellids, most of which are Mesozoic. The detailed nature of the asymmetry is very variable, but does not extend to internal hard parts such as crura. Taken as a whole, asymmetric brachiopod species show no preference for any particular environment or geographic region, and in no circumstances seem to have been markedly more or less successful than symmetric species. We are thus led to suggest that asymmetry was a genetically based condition which cropped up periodically in brachiopod evolution, and which possibly was selected neither particularly for nor against. 0 Brachiopoda, asymmetry, functional morphology, mode of life.
Ancient marine organisms that excavated holes in shells and other calcium carbonate substrates on the sea bed have not been widely studied by palaeontologists. Together with scrape and scratch marks made by rasping grazers on shell surfaces, they constitute a group of hard–substrate trace fossils that have many modern equivalents. Even the smallest forms can be studied by casting the holes in plastic resin and dissolving the shell. The variety of borers increased in the Mesozoic Era, thereby avoiding the attentions of predatory groups that were radiating at the same time.
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