Savannas are globally important ecosystems of great significance to human economies. In these biomes, which are characterized by the co-dominance of trees and grasses, woody cover is a chief determinant of ecosystem properties. The availability of resources (water, nutrients) and disturbance regimes (fire, herbivory) are thought to be important in regulating woody cover, but perceptions differ on which of these are the primary drivers of savanna structure. Here we show, using data from 854 sites across Africa, that maximum woody cover in savannas receiving a mean annual precipitation (MAP) of less than approximately 650 mm is constrained by, and increases linearly with, MAP. These arid and semi-arid savannas may be considered 'stable' systems in which water constrains woody cover and permits grasses to coexist, while fire, herbivory and soil properties interact to reduce woody cover below the MAP-controlled upper bound. Above a MAP of approximately 650 mm, savannas are 'unstable' systems in which MAP is sufficient for woody canopy closure, and disturbances (fire, herbivory) are required for the coexistence of trees and grass. These results provide insights into the nature of African savannas and suggest that future changes in precipitation may considerably affect their distribution and dynamics.

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In the Okavango Delta (about 28,000 km 2 ) the number of identifi ed species is 1,300 for plants, 71 for fi sh, 33 for amphibians, 64 for reptiles, 444 for birds, and 122 for mammals. The local occurrence of different species of these taxonomic groups in the Okavango Delta is mainly due to a hydrological gradient from permanent streams and swamps to seasonal fl oodplains, riparian woodlands, and dry woodlands. This level of species diversity is normal for the southern African region, and all analyzed aquatic groups are composed of ubiquitous species with an additional signifi cant proportion of species originating from northern, more tropical systems. Cyclical variations in climate over thousands of years have created a huge wetland complex in the upper Zambezi and Okavango Rivers during wet phases. This wetland complex has fragmented into the Okavango Delta and other large wetlands in Zambia during dry phases. There are no endemic species in the Okavango Delta while the Southcentral African wetland complex is a centre of endemism. Species diversity of the Okavango Delta is a consequence of this unique environment, with dynamic shifts in fl ooding patterns that in turn force constant changes in patterns of plant succession and dependent animals. Temporal variations in fl ooding also cause accumulation and sudden mobilization of nutrients which are readily used by well adapted plant species. As a consequence, locally high biological productivity occurs, which in turn results in high numbers of grazing mammals.

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Changes in vegetation cover in northern Chobe National Park (Botswana) were assessed using aerial photographs from1962,1985 and1998, with subsequent ground proofing. In addition, cumulative browsing by elephants and the occurrence of ¢re scars were recorded on random vegetation sites within shrubland (n 20) and mixed woodland (n 20). Coverage of woodland vegetation decreased from 60% to 30% between 1962 and 1998, while shrubland vegetation increased from 5% to 33% during the same period. During the study period, woodland has gradually retreated away from the river front. While riparian forest covered a continuous area along the riverfront in 1962, only fragments were left in 1998. We found a signi¢cant decrease in browse use with increasing distance to the Chobe river for Combretum apiculatum, Combretum elaeagnoides, Combretum mossambicense and other woody plants combined (all P < 0.0001). The occurrence of ¢re (P < 0.0001) and basal area (P < 0.0001) were positively related to distance to the river. Elephant browsing occurred on >70% of available stems within 2 km from the river, while less than 20% of the trees had ¢re scars in the same zone. Beyond 7 km from the river, elephant browsing was reduced to >50% of available stems, while more than 50% of the trees had ¢re scars. The density of any of the shrubs was not related to distance to the river neither within shrubland (all P > 0.05) nor within mixed woodlands (all P > 0.05).La densite¨d'aucunarbuste n'e¨tait lie¨e a© la distance jusqu'a© la rivie© re, ni dans la zone arbustive (ensemble P > 0.05) ni dans la zone arbore¨e me¨lange¨e (ensemble P > 0.05).

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