Pleurotomaria species from lower Bajocian (Middle Jurassic) sediments of south-western Luxembourg housed in the National Natural History Museum of Luxembourg are described. Seven species are recognized, one of which is new, Pleurotomaria faberi sp. nov. A more detailed definition of the diagnostic characters of the genus is proposed and the morphological continuity between Talantodiscus and Pleurotomaria is demonstrated, suggesting that the former cannot be considered as a distinct taxon. The palaeoecology, evolution and palaeobiogeographical history of Pleurotomaria are outlined. Pleurotomaria presumably first appeared in late Middle Triassic of New Zealand where it underwent a relative diversification up to the Hettangian (Early Jurassic). From early Hettangian, most of its evolutionary history took place in Europe and western Tethys. In the European epicontinental seas, Pleurotomaria experienced two important radiations. The first occurred in the Early Jurassic, with a peak in the late Pliensbachian, and was marked by an expansion of the distribution to the central part of western Tethys. After a collapse in species diversity, probably related to the early Toarcian anoxic event, a second radiation occurred. This culminated in the early Bajocian and was mainly confined in a region encompassing southern England, Paris Basin and southern Germany. Low-spired species, formerly attributed to Talantodiscus, probably originated independently and iteratively during the history of Pleurotomaria. The facies and associated benthic faunas suggest that Pleurotomaria probably lived on shallow soft bottoms composed of mixed calcareous-siliciclastic sediments. The two main Early Jurassic and early Middle Jurassic radiations of the genus took place in these environments. Records of the genus in Jurassic carbonate platform deposits are very few and concern mainly post-Bajocian species.

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The Pliensbachian gastropods described by De Toni in 1912, coming from an isolated boulder at the foot of Mt Vedana (eastern margin of Trento Platform, Venetian Southern Alps, Italy) are revised. The fauna consists of 13 species representing nine families and eight superfamilies. Despite the low number of species, the assemblage represents the most diverse Early Jurassic gastropod fauna known for the Venetian Southern Alps. The boulder yielding the material was thought to derive from the upper part of the Early Jurassic Calcari Grigi Group, a carbonate platform unit extensively cropping out in the Mt Vedana area. The sedimentological analysis indicates a prevalently bioclastic wackstone-floatstone, reflecting a lime-muddy deposit undergone to an early consolidation. This and the high content of ammonoids, which is unusual for the Calcari Grigi Group, are typical aspects of a condensed pelagic sediment, presumably a fissure filling at the top of the carbonate platform succession. Palaeobiogeographical comparisons show that the fauna is composed of species occurring exclusively in pelagic limestones of the western Tethys. By contrast, it shows no relationships with the coeval faunas of the adjacent Trento platform and of the other western Tethyan carbonate platforms. These lines of evidence and the facies analysis would testify to the Pliensbachian drowning of the eastern margin of the Trento platform. In the wider context of the palaeobiogeographical history of Early Jurassic western Tethyan gastropods, the species from Vedana belong to a faunal stock which is typical for pelagic, mainly postdrowning sediments. Thus, appearance and diffusion in space and time of this stock were probably regulated by the direction, rate and pattern of the Neotethyan rifting. A new subgenus, Proarcirsa (Schafbergia) subgen. nov., and three new species, namely Ataphrus (Ataphrus) cordevolensis sp. nov., Guidonia pseudorotula sp. nov. and Proarcirsa (Schafbergia) zirettoensis sp. nov. are erected.

Vetigastropod and neritimorph species from Lower Bajocian (Middle Jurassic) sediments of south-western Luxembourg are described. Eighteen species are recognized. Two new genera – Szabotomaria gen. nov. and Fabercapulus gen. nov. – and four new species – Szabotomaria ziqquratiformis sp. nov., Colpomphalus thuyi sp. nov., Colpomphalus tigratus sp. nov. and Fabercapulus semisculptus sp. nov. – are erected. These species, together with other species recently described from the same strata, are part of a diverse assemblage consisting of 32 species belonging to 14 genera in six families and five superfamilies, representing the richest vetigastropod-neritimorph fauna currently known from Bajocian and sub-coeval deposits of western Europe. An analysis of the palaeoecological and palaeobiogeographical relationships of this fauna and those from other areas of the western European shelf was performed using cluster analysis and by comparison of the respective taxonomic structures. The former detected a major cluster composed of two distinct branches: the faunas of the northern Paris-Wessex Basin, including Luxembourg, and those of the southern Germany basin. These faunas have similar taxonomic structures and occur mainly in facies represented by condensed iron ooid-rich marls and limestones deposited in lower offshore to upper offshore-shoreface settings. The close relationship between the Luxembourg and other faunas of the Paris-Wessex Basin reflects free faunal exchange and facies similarities. Slight differences in the sedimentary context could explain the separation of the southern German from the Anglo-Paris branch, whereas similarities between Swabian and Franconian faunas most probably reflect the geographical continuity of these areas. The Early Bajocian fauna of the East Midlands Shelf comes from sediments deposited in a shallow-water, oolitic barrier complex and is different from\ud other western European faunas in both species composition and taxonomic structure

A gastropod fauna has been studied from upper Pliensbachian -upper Toarcian deposits of two sections of the Causses Basin (southern France) in order to investigate the mode of recovery after the early Toarcian anoxic event. The fauna consists of 15 species, one of which is new (Bathrotomaria kronzwilmesorum sp. nov.). Their stratigraphical distribution shows two peaks of diversity -in the Bifrons Zone (Bifrons Subzone) and in the Aalensis Zone (Mactra Subzone) -which reflect brief times during which the oxygen content and bottom consistency favoured the settlement of a relatively diversified fauna. In the Variabilis-Pseudoradiosa zones, gastropods are only represented by two species. This probably indicates more severe and unstable environmental conditions, only allowing the survival of gastropod taxa with wide adaptive capacities. The very low species diversity and the discontinuous and slow faunal recovery were probably determined by physiographic factors. The Causses area was a small basin confined by exposed lands and open towards the central part of western Tethys. Gastropods described here occur exclusively in the Toarcian -early Aalenian communities of the European epicontinental seas, whereas species from the central region of western Tethys are absent. Geographic isolation and marginal location of the Causses Basin restricted faunal exchange with the western European epicontinental seas, preventing fast recovery after the anoxic event. Gastropods of the central region of the western Tethys were probably unable to settle and colonize that area due to the strongly different environment.

ABSTRACT. A new lucinoidean genus and species, Cerkesia contiae gen. et sp. nov., from the Kimmeridgian carbonate platform deposits of the C Ë erkes area (western Pontides, Turkey), is described and placed in the family Fimbriidae. Morphological comparisons reveal similarities with Fimbria, Cyclopellatia and Sphaera. A phylogenetic analysis supports the monophyly of the family. Similarities between Cerkesia and Fimbria determine inclusion of these genera in the same clade. The unusual features characterizing Cerkesia closely relate it to Cyclopellatia. Sphaera belongs to a different clade and its similarities with Cerkesia are owing to convergent and plesiomorphic characters. Cerkesia was a slow and shallow burrower. The morphology of the anterior adductor scar suggests a mode of life comparable to that of Fimbria but other features indicate adaptive strategies typical of the Lucinidae. The associated fossils and facies reveal that Cerkesia lived in soft and sandy substrates of quiet and protected areas of the carbonate platform under stressful environmental conditions. The appearance of Cerkesia in the western Pontides marks the eastward expansion of a clade that developed principally in the European seas during the Late Jurassic while the Sphaeriola-Sphaera clade was con®ned to the southernmost Tethyan margin.