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This paper illustrates how different methodological approaches can be combined to reveal complex patterns of constructional variation and change in the diachronic development of English ing-nominals. More specifically, we argue that approaching the data from a schema-based (rather than morphemebased) perspective shows that nominal gerunds in English, from the 16 th to the 19 th century, have undergone a semantic drift towards more "nouny" construal variants. This hypothesis is supported not only by raw frequency counts, but also by association measures and by a detailed analysis of hapax legomena.

In recent years, multiple researchers working on the evolution of language have put forward the idea that the theoretical framework of usage-based approaches and Construction Grammar is highly suitable for modelling the emergence of human language from pre-linguistic or proto-linguistic communication systems. This also raises the question of whether usage-based and constructionist approaches can be integrated with the analysis of animal communication systems. In this paper, we review possible avenues where usage-based, constructionist approaches can make contact with animal communication research, which in turn also has implications for theories of language evolution. To this end, we first give an overview of key assumptions of usage-based and constructionist approaches before reviewing some key issues in animal communication research through the lens of usage-based, constructionist approaches. Specifically, we will discuss how research on alarm calls, gestural communication and symbol-trained animals can be brought into contact with usage-based, constructionist theorizing. We argue that a constructionist view of animal communication can yield new perspectives on its relation to human language, which in turn has important implications regarding the evolution of language. Importantly, this theoretical approach also generates hypotheses that have the potential of complementing and extending results from the more formalist approaches that often underlie current animal communication research.

Two of the main theoretical approaches to the evolution of language are biolinguistics and usage-based approaches. Both are often conceptualized as belonging to seemingly irreconcilable “camps.” Biolinguistic approaches assume that the ability to acquire language is based on a language-specific genetic foundation. Usage-based approaches, on the other hand, stress the importance of domain-general cognitive capacities, social cognition, and interaction. However, there have been a number of recent developments in both paradigms which suggest that biolinguistic and usage-based approaches are actually moving closer together. For example, theoretical advancements such as evo-devo and complex adaptive system theory have gained traction in the language sciences, leading to changed conceptions of issues like the relative influence of “nature” and “nurture.” In this paper, we outline points of convergence between current minimalist biolinguistic and usage-based approaches regarding four contentious issues: (1) modularity and domain specificity; (2) innateness and development; (3) cultural and biological evolution; and (4) knowledge of language and its description. We show that across both paradigms, researchers have come to increasingly embrace more complex views of these issues. They also have come to appreciate the view that biological and cultural evolution are closely intertwined, which lead to an increased amount of common ground between minimalist biolinguistics and usage-based approaches.