Penguins (Sphenisciformes) have arguably the most complete and continuous fossil record of any avian clade, offering an ever-improving understanding of penguin phylogeny, biogeography, and the evolution of wing-propelled diving. Yet, our knowledge of the precise body proportions of stem-group penguins remains poor due to a dearth of articulated specimens. Here, we describe Kairuku waewaeroa sp. nov., a new giant penguin species from the Glen Massey Formation (Whaingaroan stage, 34.6-27.3 Ma). The holotype skeleton, discovered in Kawhia Harbour, North Island, New Zealand, is one of the most complete skeletons of a giant penguin yet uncovered. Our phylogenetic analysis recovers a clade uniting the New Zealand endemics Kairuku waewaeroa, Kairuku waitaki, and Kairuku grebneffi, which is supported by synapomorphies including a stout femoral shaft and tibiotarsi with a distinctly convex medial condyle. Kairuku waewaeroa is unique among stem penguins in having elongate tibiotarsi, revealing a new long-legged stem penguin body plan. The discovery of Kairuku waewaeroa contributes yet another penguin species to an Oligocene avifauna for Zealandia that is replete with giant birds.

We describe a new species of extinct duck, Miotadorna catrionae sp. nov. (Anatidae, Tadornini, Tadorninae), based on a right humerus from the Miocene lacustrine deposits of St Bathans, Otago, New Zealand. Principal component analysis reveals that the new taxon is distinguished by its large size and relative proportions. This is the eighth and largest species of duck described from the St Bathans fossil assemblage and further underscores the global importance of this site for understanding anatid evolution.

Popular current methods for quantifying variation in biological shape are well‐suited to analyses of isolated parts (e.g. the same bone from the skeletons of many individuals). An analytical challenge exists for quantifying variation between the shapes of multiple‐part objects where each part has a different position, rotation or scale (e.g. partial or whole articulated skeletons). We investigated regularised consensus principal component analysis (RCPCA) as a multiblock method for quantifying variation in the shape of multiple‐part objects. Multiblock methods are routinely used in other big data research fields such as bioinformatics/medicine, marketing and food research, but have not been widely embraced for evolutionary biology research. We have created the new package morphoBlocks for the r programming language to make RCPCA more accessible for shape evolution research. morphoBlocks provides a complete workflow for formatting, analysing and visualising the variation between multiple‐part objects by integrating functions from a diverse range of other packages. In particular, global components produced by RCPCA provide a consensus space that we present here as a morphospace for multiple‐part objects. morphoBlocks is demonstrated with a case study of manually placed landmarks and automatically placed pseudolandmarks from the partial wing skeletons of 15 extant penguin species and five fossil penguin species. Our case study provides quantitative support for a historical hypothesis about the magnitude and mode of morphological change across the evolutionary history of penguins. RCPCA can be used to analyse two‐ or three‐dimensional datasets with 10s of landmarks, or 100s to 1,000s of semilandmarks or pseudolandmarks, from 10s to 100s of specimens comprised of two or more parts. We use morphoBlocks on a small three‐bone case study and provide a framework for applying this method to much larger studies investigating the ecological or evolutionary significance of multiple‐part objects.

The Morrison Formation has been explored for dinosaurs for more than 150 years, in particular for large sauropod skeletons to be mounted in museum exhibits around the world. Several long-term campaigns to the Jurassic West of the United States produced hundreds of specimens, ranging from isolated, fragmentary bones to nearly complete skeletons of these enormous herbivorous animals. Given the sheer number of specimens, keeping track of what is housed in which institution is paramount to study variability, taxonomy, and consequently geographic and temporal distribution of the various species and genera recognized from the Morrison Formation. In an attempt to facilitate these studies, we have compiled an online spreadsheet intended to combine all the available information on sauropod specimens from collection databases, published literature, and personal observations. These include lists of contents of the specimens, in what institution the material is housed, references mentioning, describing, figuring, providing measurements and/or 3D scans, locality data and stratigraphy, as well as other potentially useful data for research purposes. The spreadsheet is openly accessible, but editing is currently restricted to the authors of this study, in order to ensure high-quality data curation to keep the file as useful as possible.

Recent fossil discoveries from New Zealand have revealed a remarkably diverse assemblage of Paleocene stem group penguins. Here, we add to this growing record by describing nine new penguin specimens from the late Paleocene (upper Teurian local stage; 55.5–59.5 Ma) Moeraki Formation of the South Island, New Zealand. The largest specimen is assigned to a new species, Kumimanu fordycei n. sp., which may have been the largest penguin ever to have lived. Allometric regressions based on humerus length and humerus proximal width of extant penguins yield mean estimates of a live body mass in the range of 148.0 kg (95% CI: 132.5 kg–165.3 kg) and 159.7 kg (95% CI: 142.6 kg–178.8 kg), respectively, for Kumimanu fordycei. A second new species, Petradyptes stonehousei n. gen. n. sp., is represented by five specimens and was slightly larger than the extant emperor penguin Aptenodytes forsteri. Two small humeri represent an additional smaller unnamed penguin species. Parsimony and Bayesian phylogenetic analyses recover Kumimanu and Petradyptes crownward of the early Paleocene mainland NZ taxa Waimanu and Muriwaimanu, but stemward of the Chatham Island taxon Kupoupou. These analyses differ, however, in the placement of these two taxa relative to Sequiwaimanu, Crossvallia, and Kaiika. The massive size and placement of Kumimanu fordycei close to the root of the penguin tree provide additional support for a scenario in which penguins reached the upper limit of sphenisciform body size very early in their evolutionary history, while still retaining numerous plesiomorphic features of the flipper. UUID: https://zoobank.org/15b1d5b2-a5a0-4aa5-ba0a-8ef3b8461730