Food availability is an important factor in the life histories of organisms because it is often limiting and thus can affect growth, mass change, reproduction, and behaviors such as thermoregulation, locomotion, and mating. Experimental studies in natural settings allow researchers to examine the effects of food on these parameters while animals are free to behave naturally. The wide variation among organisms in energy demands and among environmental food resources suggest that responses to changes in food availability may vary among organisms. Since most supplemental feeding field experiments have been conducted on species with high energy demands, we conducted a supplemental feeding study on free-ranging, female Western diamond-backed rattlesnakes (Crotalus atrox), a species with low energy demands and infrequent reproductive investment. Snakes were offered thawed rodents 1-4 times per week. Over two active seasons, we collected data on surface activity, home range size, growth, mass change, and reproduction of supplementally fed and control snakes. Fed and control snakes did not differ in surface activity levels (proportion of time encountered above versus below ground) or home range size. Fed snakes grew and gained mass faster, and had a dramatically higher occurrence of reproduction than control snakes. Also, fed snakes were in better body condition following reproduction than snakes that were not fed. However, litter characteristics such as offspring number and size were not increased by feeding, suggesting that these characteristics may be fixed. These data experimentally demonstrate that food availability can directly impact some life history traits (i.e., growth and reproduction for C. atrox), but not others (i.e., surface activity and home range size for C. atrox). The relationship between food availability and life history traits is affected in a complex way by ecological traits and physiological constraints, and thus interspecific variation in this relationship is likely to be high.
Cities can enhance their sustainability planning by prioritizing accessibility as a performance metric for transportation planning. In this context, accessibility measures the ease of reaching opportunities and captures more characteristics of sustainable cities than the more conventionally used mobility metrics, focusing on the overall ability to move and generally recommending faster speeds to overcome distance. However, how cities can prioritize accessibility in planning is understudied. This research presents a case of Tempe, Arizona, putting accessibility into practice through a 20-min lens. Using a 20-min threshold for all modes, the city aims to promote travel for daily activities that is less reliant on private autos. The analysis presented here documents the challenges that planners face when planning for accessibility. The challenges range from technical points to decisions that need to be made about the quality of the built environment. The analysis also shows that Tempe, which is a classically suburban city of wide roads and single-family homes that was built around the automobile, is highly accessible by a 20-min metric by bicycling, walking and transit. These results suggest that planners focus on street network improvements that prioritize accessibility as part of a sustainability strategy.
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