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The quality of oocytes is closely associated with the successful production of porcine oocytes in vitro. However, the regulating mechanisms of oocytes in vitro maturation (IVM) need to be further improved, and the quality of oocyte maturation in vitro is still inferior to that of oocyte maturation in vivo. Compared with oocytes matured in vivo, the oocytes matured in vitro tend to have such problems as more high precision fertilization rate, low blastocyst formation rate, not fully unified oocyte cytoplasm and nuclei of mature in the current maturation systems (Jiao et al., 2020). Moreover, the mature quality of porcine oocytes has an essential effect on embryo development after in vitro fertilization (IVF), nuclear transfer (NT) and parthenogenetic activation (PA) (Kim et al., 2010). Oocyte

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