Although forest edges have been studied extensively as an important consequence of fragmentation, a unifying theory of edge influence has yet to be developed. Our objective was to take steps toward the development of such a theory by (1) synthesizing the current knowledge of patterns of forest structure and composition at anthropogenically created forest edges, (2) developing hypotheses about the magnitude and distance of edge influence that consider the ecological processes influencing these patterns, and (3) identifying needs for future research. We compiled data from 44 published studies on edge influence on forest structure and composition in boreal, temperate, and tropical forests. Abiotic and biotic gradients near created forest edges generate a set of primary responses to edge creation. Indirect effects from these primary responses and the original edge gradient perpetuate edge influence, leading to secondary responses. Further changes in vegetation affect the edge environment, resulting in ongoing edge dynamics. We suggest that the magnitude and distance of edge influence are a direct function of the contrast in structure and composition between adjacent communities on either side of the edge. Local factors such as climate, edge characteristics, stand attributes, and biotic factors affect patch contrast. Regional factors define the context within which to assess the ecological significance of edge influence (the degree to which the edge habitat differs from interior forest habitat). Our hypotheses will help predict edge influence on structure and composition in forested ecosystems, an important consideration for conservation. For future research on forest edges in fragmented landscapes, we encourage the testing of our hypotheses, the use of standardized methodology, complete descriptions of study sites, studies on other types of edges, synthesis of edge influence on different components of the ecosystem, and investigations of edges in a landscape context. Harper et al. Edge Influence in Fragmented Landscapes 769 Resumen: Aunque los bordes de bosque han sido extensivamente estudiados como una importante consecuencia de la fragmentación, aún no se ha desarrollado una teoría unificadora de la influencia del borde. Nuestros objetivos fueron acercarnos al desarrollo de tal teoría mediante (1) la síntesis del conocimiento actual de los patrones de estructura y composición del bosque en bordes de bosque creados antropogénicamente; (2) el desarrollo de hipótesis relacionadas con la magnitud y distancia de la influencia del borde considerando procesos ecológicos que influyen sobre esos patrones; y (3) la identificación de futuras necesidades de investigación. Recopilamos datos de 44 estudios publicados sobre la influencia del borde sobre la estructura y composición de bosques boreales, templados y tropicales. Los gradientes bióticos y abióticos creados cerca de los bordes de bosque generan una serie de respuestas primarias a la creación del borde. Los efectos indirectos de estas respuestas primarias y el borde ori...

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We studied canopy gaps over a range of stand ages and site moisture classes in the temperate rain forest of Clayoquot Sound, southwestern British Columbia, Canada. We tested predictions about the landscape mosiac at three levels of resolution: the forest as a whole, gaps, and gapmakers. In addition to canopy gaps arising from the patchy mortality of dominant trees (developmental gaps), we described the prevalence and characteristics of gaps arising from edaphic and topographic features. Roughly 56% of the forest ares is influenced to some degree by canopy openings (canopy gap + expanded gap), with 30% of forest area in canopy gaps averaged over all sites (14% developmental canopy gaps and 16% in edaphic canopy gaps). Within the most intensively sampled biogeoclimatic subzone, 73% of gaps were developmental in origin. The majority of edapthic gaps result from streamcourses. Though percent area in developmental canopy gaps was similar across stand age classes, gaps and gapmaker were more common in old growth than in mature stands. Most developmental gaps resulted from the mortality of more than one tree: 96% of the developmental gaps had more than one gapmaker, and 50% had three or more. Mature stands had a significantly higher number of gapmakers per gap than old—growth stands. Old—growth forests were thus dominated by many small gaps and mature forests by fewer larger gaps. Drier and wetter sites had more gapmakers per gap than did stands on mesic sites. The number of decay classes of gapmakers in developmental gaps increases with number of gapmakers per gap, indicating that larger gaps represent the combined effects of distinct mortality events separated in time, rather than single large events. Gapmakers in developmental gaps were distributed broadly and evenly over the range of decay classes, suggesting a continuous pattern of recruitment of gapmakers rather than larger episodic mortality events. The snapping of boles and standing death were the most common modes of gapmakers mortality, together accounting for 76% of the mortality of gapmakers. Uprooting was the least common mode of gapmaker mortality in both mature and old—growth forests (23.6% and 15.6%, respectively; 20.6% of all gapmakers) and uprooting was more common in drier and wetter sites than in mesic sites. These general trends in gap—phase structure and gap formation processes are consistent with data on gaps and forests age structure collected elsewhere on the British Columbia coast, suggesting that a regime of small—scale, low—intensity disturbance is common in the old—growth forests of coastal British Columbia. We estimate that, in the absence of large—scale disturbances, turnover time for this forest is between 350 and 950 yr, resulting in ubiquitous late successional character throughout the landscape mosaic. Silvicultural practices intended to emulate the natural disturbance regime of these forests should create small gaps of 3—10 trees in an otherwise continuous forest matrix and should retain substantial late—successional characteristics wi...

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